Uggla drove in two runs and has seven RBIs and three homers in two games; he hit a grand slam Saturday and has 11 homers this season.
The Marlins have won all six of their games at Nationals Park, which opened this season, and their seven-game win streak overall is their best run since winning nine straight in August 2006.
Washington's Aaron Boone had a homer, triple and single with two RBIs in the loss.
Logan Kensing (2-0) pitched a scoreless seventh inning for the win, and Kevin Gregg pitched the ninth for his seventh save in eight chances.
Luis Ayala (1-3) gave up three runs on two homers in the eighth inning, spoiling Shawn Hill's solid outing.
Hill allowed two runs on five hits and struck out three in seven innings, but got a no-decision for the fifth time in five starts this season.
Alfredo Amezaga led off the eighth against Ayala with an infield single. The next batter, Jeremy Hermida hit a tying two-run home run to right-center.
Then with two outs, Uggla put the Marlins ahead with a shot over the out-of-town scoreboard in right field. Uggla homered to left-center in the fifth inning.
Uggla scored the Marlins' first run in the second inning. After being hit by a pitch, he advanced to third on a bloop single by Luis Gonzalez, and then came home on Wes Helm's sacrifice fly to center.
Boone tied the game at 1 in the bottom of the inning with a leadoff home run.
The Nationals took their first lead of the three-game series in the third inning. Ronnie Belliard hit a two-run home run, his second of the season, to give Washington a 3-1 lead. Ryan Zimmerman followed with an infield single and then scored when Boone tripled.
[Associated Press]
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